I have confirmed this as C. atrovespa because this is the currently accepted name for the local Green-comb Spider Orchid, although recognising that it is very closely related to C. tentaculata. For a lot of the Spider orchids the only way of separating them is by the pollinator!
This may well be Caladenia atrovespa - I think ON AVERAGE they have longer petals and sepals but there is much overlap in most traits with Caladenia phaeoclavia. They are definitely different species, they emit different chemical scents and thereby attract different wasp pollinators. Matt - natural selection can definitely lead to different populations evolving different so-called semiochemicals and thereby being pollinated by different pollinators. This process can even happen within orchid populations. As to when things are considered separate species, that is an old and tricky question. Usually species are considered distinct when there is no gene flow between them i.e. there is generally no hybridisation. Obviously there are exceptions to this, but the sexually deceptive spider orchids have extremely strong pre-pollinating barriers (i.e. pollen from C. atrovespa never ends up on C. phaeoclavia plants, because they use two very distinct pollinators). As far as I know so far, there is no way to definitively tell C. atrovespa from C. phaeoclavia, apart from to see which pollinator they attract - not very helpful I know!
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